These studies identified four ancestral taxons
C. medica. (citron),
C.reticulata Blanco (mandarin),
C.maxima (Burm.) Merr. (pummelo), and
C.micranthaWester (papeda)]
as the ancestors of all cultivated Citrus

.../ Toutes les adhésions de sinensis de C., d'aurantium de C., de C. paradisi, d'hybrides de mandarine, de tangors, de tangelos, et d'orangelo ont eu une structure héréditaire de mélange de deux taxons (C. reticulata de C. et maxima ) avec des contributions variables.
.../A 454 amplicon haplotype study for chromosome 2 [ 51 ] also revealed introgression by C . maxima in nine of the thirteen mandarins studied. In the present work, species-diagnostic marker analysis revealed such C . maxima introgression in 13 of the 17 mandarin analysed. Only three mandarins were found without any indication of interspecifi c introgression ( ‘ Cleopatra ’ , ‘ Sunki ’ , and C . daoxianensis ). C . daoxianensis is considered as an ancestral mandarin [ 67 ], and the two others are mostly used as rootstock